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Monday, February 24, 2020
Eastern Time: 6:06 PM || Central Time: 5:06 PM || Mountain Time: 4:06 PM || Pacific Time: 3:06 PM


Amazon River Dolphin

Click the blue ball to see Pictures...

Inia geoffrensis

Also known as the Boto or pink river dolphin, is a species of toothed whale classified in the family Iniidae. Three subspecies are currently recognized:

1. I. g. geoffrensis (Amazon river dolphin)
2. I. g. boliviensis (Bolivian river dolphin)
3. I. g. humboldtiana (Orinoco river dolphin).

The three subspecies are distributed in the Amazon basin, the upper Madeira River in Bolivia, and the Orinoco basin, respectively.

Size: The Amazon river dolphin is the largest species of river dolphin, with adult males reaching 408 lb in weight, and 8.2 ft in length. Adults acquire a pink color, more prominent in males, giving it its nickname "pink river dolphin". Sexual dimorphism is very evident, with males measuring 16% longer and weighing 55% more than females. Like other toothed whales, they have a melon, an organ that is used for bio sonar.

Fins: The dorsal fin, although short in height, is regarded as long, and the pectoral fins are also large. The fin size, unfused vertebrae, and its relative size allow for improved manoverability when navigating flooded forests and capturing prey.

Range: They have one of the widest ranging diets among toothed whales.

Feeding: and feed on up to 53 different species of fish, such as croakers, catfish, tetras and piranhas. They also consume other animals such as river turtles and freshwater crabs.

Protected?: In 2008, this species was ranked by the International Union for Conservation of Nature (IUCN) as a Data Deficient, due to the uncertainty regarding their population trends and the impact of threats. While hunting is a major threat, in recent decades, greater impacts on population are due to the loss of habitat and inadvertent entanglement in fishing lines. It is the only species of river dolphin kept in captivity, mainly in the United States, Venezuela and Europe; however, it is difficult to train and a high mortality is seen in captive species.

Re-classified: The species Inia geoffrensis was described by Henri Marie Ducrotay de Blainville in 1817. Originally, the Amazon river dolphin belonged to the superfamily Platanistoidea, which constituted all river dolphins, making them a paraphyletic group. Today, however, the Amazon river dolphin has been reclassified into the superfamily Inioidea. There is no consensus on when and how they penetrated the Amazon basin; they may have done so during the Miocene from the Pacific Ocean, before the formation of the Andes, or from the Atlantic Ocean.

Three subspecies are recognized:
1. I. g. geoffrensis (Araguaian river dolphin).
2. I. g. boliviensis (Bolivian river dolphin).
3. I. g. humboldtiana (Orinoco river dolphin).

However, based on skull morphology in 1994, it was proposed that I. g. boliviensis was a different species. In 2002, following the analysis of mitochondrial DNA specimens from the Orinoco basin, the Putumayo River (tributary of the Amazon) and the Tijamuchy and Ipurupuru rivers, geneticists proposed that genus Inia be divided into at least two evolutionary lineages: one restricted to the river basins of Bolivia and the other widely distributed in the Orinoco and Amazon. A 2014 study supports the notion of the Bolivian group as a distinct species and identifies a third species in the Araguaia-Tocantins basin.

Sub-species 1: Inia geoffrensis geoffrensis inhabits most of the Amazon River, including rivers Tocantins, Araguaia , low Xingu and Tapajos, the Madeira to the rapids of Porto Velho, and rivers Purus, Yurua, Ica, Caqueta, Branco, and the Rio Negro through the channel of Casiquiare to San Fernando de Atabapo in the Orinoco river, including its tributary: the Guaviare.

Sub-species 2: Inia humboldtiana geoffrensis are located in the Orinoco River basin, including the Apure and Meta rivers. This subspecies is restricted, at least during the dry season, to the waterfalls of Rio Negro rapids in the Orinoco between Samariapo and Puerto Ayacucho, and the Casiquiare canal.

Sub-species 3: Inia geoffrensis boliviensis has populations in the upper reaches of the Madeira River, upstream of the rapids of Teotonio, in Bolivia. It is confined to the Mamore River and its main tributary, the Iténez.

Size: The Amazon river dolphin is the largest river dolphin. Adult males reach a maximum length and weight of 8.4 ft the average 7.6 ft and 408 lb and the average 340 lb, while females reach a length and weight of 7.1 ft and 330 lb and the average 220 lb. It has very evident sexual dimorphism, with males measuring and weighing between 16% and 55% more than females, making it unique among cetaceans, where females are generally larger than males.

Coloring: The body color varies with age. Newborns and the young have a dark grey tint, which in adolescence transforms into light grey, and in adults turns pink as a result of repeated abrasion of the skin surface. Males tend to be pinker than females due to more frequent trauma from intra-species aggression. The color of adults varies between solid and mottled pink and in some adults the dorsal surface is darker. It is believed that the difference in color depends on the temperature, water transparency, and geographical location. There is one albino on record, kept in an aquarium in Germany.

Lifespan: Life expectancy of the Amazon river dolphin in the wild is unknown, but in captivity, the longevity of healthy individuals has been recorded at between ten and thirty years. However, the average longevity in captive animals is only 33 months. An individual named Apure at Duisburg Zoo, Germany, is estimated to have lived more than forty years, being captured from the wild at age nine.

Behavior: The Amazon river dolphin tends to be solitary but is sometimes seen in groups. Pods usually contain up to four individuals. Typically, social bonds occur between mother and child, but may also been seen in heterogeneous groups or bachelor groups. The largest congregations are seen in areas with abundant food, and at the mouths of rivers. There is significant segregation during the rainy season, with males occupying the river channels, while females and their offspring are located in flooded areas. However, in the dry season, there is no such separation.

Captive vs Free: Captive studies have shown that the Amazon river dolphin is less shy than the bottlenose dolphin, but also less sociable. It is very curious and has a remarkable lack of fear of foreign objects. However, dolphins in captivity may not show the same behavior that they do in their natural environment, where they have been reported to hold the oars of the fishermen, rub against the boat, pluck underwater plants, and play with sticks, logs, clay, turtles, snakes, and fish.

Courtship: Adult males have been observed carrying objects in their mouths such as branches or other floating vegetation, or balls of hardened clay. The males appear to carry these objects as a socio-sexual display which is part of their mating system. The behavior is "triggered by an unusually large number of adult males and/or adult females in a group, or perhaps it attracts such into the group. A plausible explanation of the results is that object carrying is aimed at females and is stimulated by the number of females in the group, while aggression is aimed at other adult males and is stimulated by object carrying in the group." Before determining that the species had an evident sexual dimorphism, it was postulated that the river dolphins were monogamous. Later, it was shown that males were larger than females and are documented wielding an aggressive sexual behavior in the wild and in captivity. Males often have a significant degree of damage in the dorsal, caudal, and pectoral fins, as well as the blowhole, due to bites and abrasions. They also commonly have numerous secondary teeth-raking scars. This suggests fierce competition for access to females, with a polygynous mating system, though polyandry and promiscuity cannot be excluded.

In captivity, courtship and mating foreplay have been documented. The male takes the initiative by nibbling the fins of the female, but reacts aggressively if the female is not receptive. A high frequency of copulations in a couple was observed; they used three different positions: contacting the womb at right angles, lying head to head, or head to tail.

Reproduction: Breeding is seasonal, and births occur between May and June. The period of birthing coincides with the flood season, and this may provide an advantage because the females and their offspring remain in flooded areas longer than males. As the water level begins to decrease, the density of food sources in flooded areas increases due to loss of space, providing enough energy for infants to meet the high demands required for growth.

Gestation: Is estimated to be around eleven months and captive births take 4 to 5 hours. At birth, calves are 31 inches long and in captivity have registered a growth of 0.69 ft per year. Lactation takes about a year. The interval between births is estimated between 15 and 36 months, and the young dolphins are thought to become independent within two to three years.

Mother and calves: The relatively long duration of breastfeeding and parenting suggests a strong mother-child bond. Most couples observed in their natural environment consist of a female and her calf. This suggests that long periods of parental care contribute to learning and development of the young.

Food: The diet of the Amazon river dolphin is the most diverse of the toothed whales. It consists of at least 53 different species of fish, grouped in 19 families. The prey size is between 2.0 and 31.5 inches with an average of 7.9 inches. The most frequently consumed fish belong to the families Sciaenidae (croakers), Cichlidae, and Characidae (tetras and piranhas). The dolphins dentition allows it to access shells of river turtles and freshwater crabs. The diet is more diverse during the wet season, when fish are spread in flooded areas outside riverbeds, thus becoming more difficult to catch. The diet becomes more selective during the dry season when prey density is greater.

[Dolphins and boats: Usually, these dolphins are active and feeding throughout the day and night. However, they are predominantly crepuscular. They consume about 5.5% of their body weight per day. They sometimes take advantage of the disturbances made by boats to catch disoriented prey. Sometimes, they associate with the distantly-related tucuxi (Sotalia fluviatilis), and giant otters (Pteronura brasiliensis) to hunt in a coordinated manner, by gathering and attacking fish stocks at the same time. Apparently, there is little competition for food between these species, as each prefers different prey. It has also been observed that captive dolphins share food.

Calls: Like other dolphins, river dolphins use whistling tones to communicate. The issuance of these sounds is related to the time they return to the surface before diving, suggesting a link to food. Acoustic analysis revealed that the vocalisations are different in structure from the typical whistles of other species of dolphins.

Distribution: The main branch of the Amazon River near Fonte Boa, Brazil, with multiple floodplains, lakes and smaller channels. The Amazon river dolphin is observed here throughout the year.

Habitat: Amazon river dolphins are the most widespread river dolphins. They are present in six countries in South America: Bolivia, Brazil, Colombia, Ecuador, Peru, and Venezuela, in an area covering about 2,700,000 sq miles. The boundaries are set by waterfalls, such as the Xingu and Tapajós rivers in Brazil, as well as very shallow water. A series of rapids and waterfalls in the Madeira River have isolated one population, recognized as the subspecies I. g. boliviensis, south of the Amazon basin in Bolivia.

They are also distributed in the basin of the Orinoco River, except the Caroni River and the upper Caura River in Venezuela. The only connection between the Orinoco and the Amazon is through the Casiquiare canal. The distribution of dolphins in the rivers and surrounding areas depends on the time of year; in the dry season they are located in the river beds, but in the rainy season, when the rivers overflow, they disperse to the flooded areas, both the forests and the plains.

The Amazon river dolphin is located in most of the areas aquatic habitats, including; river basins, major courses of rivers, canals, river tributaries, lakes, and at the ends of rapids and waterfalls. Cyclical changes in the water levels of rivers take place throughout the year. During the dry season, dolphins occupy the main river channels, and during the rainy season, they can move easily to smaller tributaries, to the forest, and to floodplains.

Males and females appear to have selective habitat preferences, with the males returning to the main river channels when water levels are still high, while the females and their offspring remain in the flooded areas as long as possible; probably because it decreases the risk of aggression by males toward the young and predation by other species.

The Amazon river dolphin has historically been kept in dolphinariums. Today, only three exist in captivity: one in Acuario de Valencia in Venezuela, one in Zoologico de Guistochoca in Peru, and one in Duisburg Zoo in Germany. Several hundred were captured between the 1950s and 1970s, and were distributed in dolphinariums throughout the US, Europe, and Japan. Around 100 went to US dolphinariums, and of that, only 20 survived; the last died at the Pittsburgh Zoo in 2002.

Threats: The region of the Amazon in Brazil has an extension of 1,900,000 sq miles containing diverse fundamental ecosystems. One of these ecosystems is a floodplain, or a várzea forest, and is home to a large number of fish species which are an essential resource for human consumption. The várzea is also a major source of income through excessive local commercialized fishing. Várzea consists of muddy river waters containing a vast number and diversity of nutrient rich species. The abundance of distinct fish species lures the Amazon River dolphin into the várzea areas of high water occurrences during the seasonal flooding.

Predators: In addition to attracting predators such as the Amazon river dolphin, these high-water occurrences are an ideal location to draw in the local fisheries. Human fishing activities directly compete with the dolphins for the same fish species, the tambaqui (Colossoma macropomum) and the pirapitinga (Piaractus brachypomus), resulting in deliberate or unintentional catches of the Amazon river dolphin. The local fishermen over fish and when the Amazon River dolphins remove the commercial catch from the nets and lines, it causes damages to the equipment and the capture, as well as generating ill will from the local fishermen. The negative reactions of the local fishermen are also attributed to the Brazilian Institute of Environment and Renewable Natural Resources prohibition on killing the Amazon river dolphin, yet not compensating the fishermen for the damage done to their equipment and catch.

During the process of catching the commercialized fish, the Amazon river dolphins get caught in the nets and exhaust themselves until they die, or the local fishermen deliberately kill the entangled dolphins. The carcasses are discarded, consumed, or used as bait to attract a scavenger catfish, the piracatinga (Calophysus macropterus). The use of the Amazon river dolphin carcass as bait for the piracatinga dates back to 2000. Increasing demand for the piracatinga has created a market for distribution of the Amazon river dolphin carcasses to be used as bait throughout these regions.

Of the 15 dolphin carcasses found in the Japurá River in 2010–2011 surveys, 73% of the dolphins were killed for bait, disposed of, or abandoned in entangled gillnets. The data do not fully represent the actual overall number of deaths of the Amazon river dolphins, whether accidental or intentional, because a variety of factors make it extremely complicated to record and medically examine all the carcasses. Scavenger species feed upon the carcasses, and the complexity of the river currents make it nearly impossible to locate all of the dead animals. More importantly, the local fishermen do not report these deaths out of fear that a legal course of action will be taken against them, as the Amazon river dolphin and other cetaceans are protected under a Brazilian federal law prohibiting any takes, harassments, and kills of the species.

Conservation: In 2008, the International Whaling Commission (IWC) expressed concern for captured botos for use as bait in the Central Amazon, which is an emerging problem that has spread on a large scale. The species is listed in Appendix II of the Convention on International Trade in Endangered Species Fauna and Flora (CITES), and Appendix II of the Convention on the Conservation of Migratory Species of Wild Animals, because it has an unfavorable conservation status or would benefit significantly from international co-operation organized by tailored agreements.

According to a previous assessment by the Scientific Committee of the International Whaling Commission in 2000, the population of botos appears great and there is little or no evidence of population decline in numbers and range. However, increased human intervention on their habitat is expected to, in the future, be the most likely cause of the decline of its range and population. A series of recommendations were issued to ensure proper follow-up to the species, among which is the implementation and publication of studies on the structure of populations, making a record of the distribution of the species, information about potential threats as the magnitude of fishing operations and location of pipelines.

In September 2012, Bolivian President Evo Morales enacted a law to protect the dolphin and declared it a national treasure.

In 2008, the species was listed on the Red list of endangered species, but in 2011, the IUCN stated it as Data Deficient (DD). The species was previously listed as "vulnerable" but the conservation status changed due to the limited amount of currently available information on threats, ecology, and population trends. In areas where these dolphins have been studied, they appear well extended and relatively abundant. However, these areas represent only a small proportion of the total distribution of the species and are often sites where the animals are protected. Consequently, the information from these areas may not be representative, and may not be valid in the long term.

This page was Last Updated on Wed Oct 12, 2016 2:10 pm CDT by Ladywildlife

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